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Cantabria Labs type material holotype spain cantabria province • ♂
Geographic distribution of Ischyropsalis impressa sp. nov. (yellow dots) and I. aguerana sp. nov. (white dots) in northeastern Cantabrian Mountains. A) Location of study area and map of the Karst in <t>Spain</t> at 1:1,000,000-scale where karstified carbonate rocks emerge. B) LiDAR visualisation of the study area <t>showing</t> <t>Cantabria–Biscay</t> border (solid yellow line). The orange-brown area represents the range of limestone outcrops from the Urgonian formation (of Aptian age, Early Cretaceous) at 1:50,000-scale. Location of the caves surveyed: 1. La Baja; 2. Hoyo de Peñaflor; 3. Perelada; 4. Hoyo Molino; 5. Hoyo Menor-Tocinos; 6. Hoyo Menor-Rejuyo; 7. Hoyo de la Cubilla; 8. La Lastrilla; 9. Covacho del Cogorio; 10. Hoyo de las Rebolligas; 11. La Pasada; 12. Hoyo de las Fuentes; 13. La Tabla II; 14. Gueriza.
Type Material Holotype Spain Cantabria Province • ♂, supplied by Cantabria Labs, used in various techniques. Bioz Stars score: 86/100, based on 1 PubMed citations. ZERO BIAS - scores, article reviews, protocol conditions and more
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Cientifica Ltd holotype ♂
Geographic distribution of Ischyropsalis impressa sp. nov. (yellow dots) and I. aguerana sp. nov. (white dots) in northeastern Cantabrian Mountains. A) Location of study area and map of the Karst in <t>Spain</t> at 1:1,000,000-scale where karstified carbonate rocks emerge. B) LiDAR visualisation of the study area <t>showing</t> <t>Cantabria–Biscay</t> border (solid yellow line). The orange-brown area represents the range of limestone outcrops from the Urgonian formation (of Aptian age, Early Cretaceous) at 1:50,000-scale. Location of the caves surveyed: 1. La Baja; 2. Hoyo de Peñaflor; 3. Perelada; 4. Hoyo Molino; 5. Hoyo Menor-Tocinos; 6. Hoyo Menor-Rejuyo; 7. Hoyo de la Cubilla; 8. La Lastrilla; 9. Covacho del Cogorio; 10. Hoyo de las Rebolligas; 11. La Pasada; 12. Hoyo de las Fuentes; 13. La Tabla II; 14. Gueriza.
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Pasteur Institute ♂ holotype
Geographic distribution of Ischyropsalis impressa sp. nov. (yellow dots) and I. aguerana sp. nov. (white dots) in northeastern Cantabrian Mountains. A) Location of study area and map of the Karst in <t>Spain</t> at 1:1,000,000-scale where karstified carbonate rocks emerge. B) LiDAR visualisation of the study area <t>showing</t> <t>Cantabria–Biscay</t> border (solid yellow line). The orange-brown area represents the range of limestone outcrops from the Urgonian formation (of Aptian age, Early Cretaceous) at 1:50,000-scale. Location of the caves surveyed: 1. La Baja; 2. Hoyo de Peñaflor; 3. Perelada; 4. Hoyo Molino; 5. Hoyo Menor-Tocinos; 6. Hoyo Menor-Rejuyo; 7. Hoyo de la Cubilla; 8. La Lastrilla; 9. Covacho del Cogorio; 10. Hoyo de las Rebolligas; 11. La Pasada; 12. Hoyo de las Fuentes; 13. La Tabla II; 14. Gueriza.
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Salvin Dental Specialties holotype ♂
Geographic distribution of Ischyropsalis impressa sp. nov. (yellow dots) and I. aguerana sp. nov. (white dots) in northeastern Cantabrian Mountains. A) Location of study area and map of the Karst in <t>Spain</t> at 1:1,000,000-scale where karstified carbonate rocks emerge. B) LiDAR visualisation of the study area <t>showing</t> <t>Cantabria–Biscay</t> border (solid yellow line). The orange-brown area represents the range of limestone outcrops from the Urgonian formation (of Aptian age, Early Cretaceous) at 1:50,000-scale. Location of the caves surveyed: 1. La Baja; 2. Hoyo de Peñaflor; 3. Perelada; 4. Hoyo Molino; 5. Hoyo Menor-Tocinos; 6. Hoyo Menor-Rejuyo; 7. Hoyo de la Cubilla; 8. La Lastrilla; 9. Covacho del Cogorio; 10. Hoyo de las Rebolligas; 11. La Pasada; 12. Hoyo de las Fuentes; 13. La Tabla II; 14. Gueriza.
Holotype ♂, supplied by Salvin Dental Specialties, used in various techniques. Bioz Stars score: 90/100, based on 1 PubMed citations. ZERO BIAS - scores, article reviews, protocol conditions and more
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Khanh Hoa Salanganes Nest Company holotype ♂
Geographic distribution of Ischyropsalis impressa sp. nov. (yellow dots) and I. aguerana sp. nov. (white dots) in northeastern Cantabrian Mountains. A) Location of study area and map of the Karst in <t>Spain</t> at 1:1,000,000-scale where karstified carbonate rocks emerge. B) LiDAR visualisation of the study area <t>showing</t> <t>Cantabria–Biscay</t> border (solid yellow line). The orange-brown area represents the range of limestone outcrops from the Urgonian formation (of Aptian age, Early Cretaceous) at 1:50,000-scale. Location of the caves surveyed: 1. La Baja; 2. Hoyo de Peñaflor; 3. Perelada; 4. Hoyo Molino; 5. Hoyo Menor-Tocinos; 6. Hoyo Menor-Rejuyo; 7. Hoyo de la Cubilla; 8. La Lastrilla; 9. Covacho del Cogorio; 10. Hoyo de las Rebolligas; 11. La Pasada; 12. Hoyo de las Fuentes; 13. La Tabla II; 14. Gueriza.
Holotype ♂, supplied by Khanh Hoa Salanganes Nest Company, used in various techniques. Bioz Stars score: 90/100, based on 1 PubMed citations. ZERO BIAS - scores, article reviews, protocol conditions and more
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Santa Cruz Biotechnology bolivia; ♂; “m. bolivianus guzman and żyła 2023 holotype [red label] \ bol/nov2013/09
Geographic distribution of Ischyropsalis impressa sp. nov. (yellow dots) and I. aguerana sp. nov. (white dots) in northeastern Cantabrian Mountains. A) Location of study area and map of the Karst in <t>Spain</t> at 1:1,000,000-scale where karstified carbonate rocks emerge. B) LiDAR visualisation of the study area <t>showing</t> <t>Cantabria–Biscay</t> border (solid yellow line). The orange-brown area represents the range of limestone outcrops from the Urgonian formation (of Aptian age, Early Cretaceous) at 1:50,000-scale. Location of the caves surveyed: 1. La Baja; 2. Hoyo de Peñaflor; 3. Perelada; 4. Hoyo Molino; 5. Hoyo Menor-Tocinos; 6. Hoyo Menor-Rejuyo; 7. Hoyo de la Cubilla; 8. La Lastrilla; 9. Covacho del Cogorio; 10. Hoyo de las Rebolligas; 11. La Pasada; 12. Hoyo de las Fuentes; 13. La Tabla II; 14. Gueriza.
Bolivia; ♂; “M. Bolivianus Guzman And żyła 2023 Holotype [Red Label] \ Bol/Nov2013/09, supplied by Santa Cruz Biotechnology, used in various techniques. Bioz Stars score: 90/100, based on 1 PubMed citations. ZERO BIAS - scores, article reviews, protocol conditions and more
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bolivia; ♂; “m. bolivianus guzman and żyła 2023 holotype [red label] \ bol/nov2013/09 - by Bioz Stars, 2026-10
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Niitaka Co Ltd holotype ♂
Geographic distribution of Ischyropsalis impressa sp. nov. (yellow dots) and I. aguerana sp. nov. (white dots) in northeastern Cantabrian Mountains. A) Location of study area and map of the Karst in <t>Spain</t> at 1:1,000,000-scale where karstified carbonate rocks emerge. B) LiDAR visualisation of the study area <t>showing</t> <t>Cantabria–Biscay</t> border (solid yellow line). The orange-brown area represents the range of limestone outcrops from the Urgonian formation (of Aptian age, Early Cretaceous) at 1:50,000-scale. Location of the caves surveyed: 1. La Baja; 2. Hoyo de Peñaflor; 3. Perelada; 4. Hoyo Molino; 5. Hoyo Menor-Tocinos; 6. Hoyo Menor-Rejuyo; 7. Hoyo de la Cubilla; 8. La Lastrilla; 9. Covacho del Cogorio; 10. Hoyo de las Rebolligas; 11. La Pasada; 12. Hoyo de las Fuentes; 13. La Tabla II; 14. Gueriza.
Holotype ♂, supplied by Niitaka Co Ltd, used in various techniques. Bioz Stars score: 90/100, based on 1 PubMed citations. ZERO BIAS - scores, article reviews, protocol conditions and more
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Arisan Inc holotype ♂
Geographic distribution of Ischyropsalis impressa sp. nov. (yellow dots) and I. aguerana sp. nov. (white dots) in northeastern Cantabrian Mountains. A) Location of study area and map of the Karst in <t>Spain</t> at 1:1,000,000-scale where karstified carbonate rocks emerge. B) LiDAR visualisation of the study area <t>showing</t> <t>Cantabria–Biscay</t> border (solid yellow line). The orange-brown area represents the range of limestone outcrops from the Urgonian formation (of Aptian age, Early Cretaceous) at 1:50,000-scale. Location of the caves surveyed: 1. La Baja; 2. Hoyo de Peñaflor; 3. Perelada; 4. Hoyo Molino; 5. Hoyo Menor-Tocinos; 6. Hoyo Menor-Rejuyo; 7. Hoyo de la Cubilla; 8. La Lastrilla; 9. Covacho del Cogorio; 10. Hoyo de las Rebolligas; 11. La Pasada; 12. Hoyo de las Fuentes; 13. La Tabla II; 14. Gueriza.
Holotype ♂, supplied by Arisan Inc, used in various techniques. Bioz Stars score: 90/100, based on 1 PubMed citations. ZERO BIAS - scores, article reviews, protocol conditions and more
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Geographic distribution of Ischyropsalis impressa sp. nov. (yellow dots) and I. aguerana sp. nov. (white dots) in northeastern Cantabrian Mountains. A) Location of study area and map of the Karst in Spain at 1:1,000,000-scale where karstified carbonate rocks emerge. B) LiDAR visualisation of the study area showing Cantabria–Biscay border (solid yellow line). The orange-brown area represents the range of limestone outcrops from the Urgonian formation (of Aptian age, Early Cretaceous) at 1:50,000-scale. Location of the caves surveyed: 1. La Baja; 2. Hoyo de Peñaflor; 3. Perelada; 4. Hoyo Molino; 5. Hoyo Menor-Tocinos; 6. Hoyo Menor-Rejuyo; 7. Hoyo de la Cubilla; 8. La Lastrilla; 9. Covacho del Cogorio; 10. Hoyo de las Rebolligas; 11. La Pasada; 12. Hoyo de las Fuentes; 13. La Tabla II; 14. Gueriza.

Journal: bioRxiv

Article Title: Unraveling the Phylogenetic Complexities of Ischyropsalis C.L. Koch, 1839 (Opiliones: Dyspnoi: Ischyropsalididae) in the North Iberian Peninsula, with the description of three new species

doi: 10.64898/2026.01.09.698449

Figure Lengend Snippet: Geographic distribution of Ischyropsalis impressa sp. nov. (yellow dots) and I. aguerana sp. nov. (white dots) in northeastern Cantabrian Mountains. A) Location of study area and map of the Karst in Spain at 1:1,000,000-scale where karstified carbonate rocks emerge. B) LiDAR visualisation of the study area showing Cantabria–Biscay border (solid yellow line). The orange-brown area represents the range of limestone outcrops from the Urgonian formation (of Aptian age, Early Cretaceous) at 1:50,000-scale. Location of the caves surveyed: 1. La Baja; 2. Hoyo de Peñaflor; 3. Perelada; 4. Hoyo Molino; 5. Hoyo Menor-Tocinos; 6. Hoyo Menor-Rejuyo; 7. Hoyo de la Cubilla; 8. La Lastrilla; 9. Covacho del Cogorio; 10. Hoyo de las Rebolligas; 11. La Pasada; 12. Hoyo de las Fuentes; 13. La Tabla II; 14. Gueriza.

Article Snippet: Type material Holotype SPAIN – Cantabria Province • ♂; Guriezo; La Pasada Cave; 30TVN7689794385, 440 m a.s.l.

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